A new girl, Connie, arrives at the farm and sets her sights on Billy. The manor house has been commandeered by the Americans, which excites Lady Hoxley's sister, Diana (Raquel Cassidy). John visits Joyce at the farm and Martin has an accident. Tucker is released from prison. Finch enters his pig into a county show, so it can avoid slaughter.

Following his fall, Martin is found to have detached retinas. He is also hiding an injured man. Diana tries to help Jack find his grandson and Joyce receives bad news about John. Finch tries to keep his contraband deals from Mrs Gulliver (Carolyn Pickles). Bea becomes friends with Salvatore Zappulo (Cesare Taurasi), an Italian POW, while Connie flirts with piano player, Henry (Liam Garrigan).

Bea allows Jack to meet his grandson. Connie discovers that Henry is actually a Reverend. Joyce follows Martin and finds Ulrich Keitmeiler (Benjamin Wilkin), the German airman he has been hiding. Esther is desperate for money to pay for Martin's operation, so Finch organises a boxing match with Vernon Storey (David Schofield).

A week later, Joyce is still in shock after shooting Ulrich dead. Connie helps Joseph Price (Sam Walton), a young evacuee, reunite with his sister and mother. Tucker works with Billy to improve his shooting and Jack tries to convince Bea to come to Chicago with him. After being turned down by Lady Hoxley, Esther asks Vernon for the money for Martin's operation.

Connie stands up to the overbearing Bishop in order to win Henry's respect. Finch blows his top when he discovers how Esther received the money for Martin's operation. Joyce is angry and disillusioned with the Land Army, and she forces Tucker into an admission of guilt. Lady Hoxley's sister Diana is part of Jack's plan to make a getaway with Bea and the baby. Billy's family is being ripped apart, and he seems disinterested; what is he hiding?

At Hoxley Manor Lady Ellen turns the Great Hall into a hospital with the help of the land girls and ends up being confronted by a face from the past when Dr Channing shows up. At Pasture Farm, Joyce is still waiting for news about John, who is believed to be missing in action. Finch ends up drunkenly driving into Mrs Gulliver's garden and causes havoc. A telegram ends up arriving for Joyce and it has good news inside, John is still alive.

After Iris is called to the farm by Vernon for her proficiency test, the huge threshing machine ends up throwing her to the ground. Connie ends up throwing down the gauntlet and challenges Vernon to a farmers versus land girls ploughing competition. Channing attempts to reassure Joyce that John needs reassurance from her. After Connie goes to see Henry at church she is shocked to see Danny there chatting to him.

John thinks about the flashbacks he is having of his escape from France and talks to Dr Channing about them. Joyce ends up intervening when John can't take any more. Esther has to deal with her problems and starts to think about her options. Finch ends up offering her another solution. Frank informs Ellen about the suspicious man he saw in the woods and she ends up calling the police.

Danny threatens to hurt Henry if Connie doesn't submit to his will. Ellen considers if she can trust the man she loves. As Joyce and John grow closer John struggles with guilt over his infidelity. A revelation drives Martin even further away from Esther. Iris is in mourning whilst Walter and Frank square up for a fight with disastrous consequences.

Events spiral out of control for Connie when her shady past catches up with her. Iris makes a shocking discovery, whilst John is given some crushing news and Esther decides to face her critics in the village. Meanwhile, can Ellen keep up the pretence with Richard?